browsed among the sea-weeds, pursued and preyed upon one another, and gave a new liveliness to the waters of the world. None of these were excessively big by our present standards. Few of them were more than two or three feet long, but there were exceptional forms which were as long as twenty feet.
We know nothing from geology of the ancestors of these fishes. They do not appear to be related to any of the formsthat preceded them. Zoologists have the most interesting views of their ancestry, but these they derive from the study of the development of the eggs of their still living relations, and from other sources. Apparently the ancestors of the vertebrata were soft-bodied and perhaps quite small swimming creatures who began first to develop hard parts as teeth round and about their mouths. The teeth of a skate or dogfish cover the roof and floor of its mouth and pass at the lip into the flattened toothlike scales that encase most of its body. As the fishes develop these teeth-scales in the geological record, they swim out of the hidden darkness of the past into the light, the first vertebrated animals visible in the record.
5
The Age of the Coal Swamps
The land during this Age of Fishes was apparently quite lifeless. Crags and uplands of barren rock lay under the sun and rain. There was no real soil â for as yet there were no earthworms which help to make a soil, and no plants to break up the rock particles into mould; there was no trace of moss or lichen. Life was still only in the sea. 1
Over this world of barren rock played great changes of climate. The causes of these changes of climate were very complex and they have still to be properly estimated. The changing shape of the Earth's orbit, the gradual shifting of the poles of rotation, changes in the shapes of the continents, probably even fluctuations in the warmth of the sun, now conspired to plunge great areas of the Earth's surface into long periods of cold and ice and now again for millions of years spread a warm or equable climate over this planet. 2 There seem to have been phases of great internal activity in the world's history, when in the course of a few million years accumulated upthrusts would break out in lines of volcanic eruption and upheaval and rearrange the mountain and continental outlines of the globe, increasing the depth of the sea and the height of the mountains and exaggerating the extremes of climate. And these would be followed by vast ages of comparative quiescence, when frost, rain and river would wear down the mountain heights and carry great masses of silt to fill and raise the sea bottoms and spread the seas, ever shallower and wider, over more and more of the land. There have been âhigh and deepâ ages in the world's history and âlow and levelâ ages. The reader must dismiss from his mind any idea that the surface of the Earth has been growing steadily coolersince its crust solidified. After that much cooling had been achieved, the internal temperature ceased to affect surface conditions. There are traces of periods of superabundant ice and snow, of âGlacial Agesâ, that is, even in the Azoic period.
It was only towards the close of the Age of Fishes, in a period of extensive shallow seas and lagoons, that life spread itself out in any effectual way from the waters onto the land. No doubt the earlier types of the forms that now begin to appear in great abundance had already been developing in a rare and obscure manner for many scores of millions of years. But now came their opportunity.
Plants no doubt preceded animal forms in this invasion of the land, but the animals probably followed up the plant emigration very closely. The first problem that the plant had to solve was the provision of some sustaining stiff support to hold up its fronds to the sunlight when the buoyant water was withdrawn; the second was the difficulty of getting water from the swampy ground below to the tissues of the plant, now
Rachel Haimowitz, Heidi Belleau